Wednesday, November 10, 2010

What is "Darwinism" and am I a "Darwinist"?


I don’t use the term “Darwinism” at all, any more than I would use the term “Newtonism” when referring to classical physical mechanics, “Einsteinism” to refer to relativity theory, “Bohr/Feinman/Heisenberg/Schroedingerism” to refer to quantum mechanics, or “Mendeleevianism” to refer to chemistry. What I and my colleagues (and friends) do is probably best described as “evolutionary biology”, and includes (at a bare minimum) the following:

1) the formulation and testing of a set of interconnected theories explaining the origin of biological diversity, consisting of descent with modification from common ancestors over deep geological time, describable via cladistic analysis, and supported by inference from multiple sources of empirical evidence, including comparative anatomy, biogeography, developmental biology, genomics, historical geology, and paleontology; and

2) the formulation and testing of a separate but related set of interconnected theories explaining the origin and modification of the phenotypic characteristics of living organisms, consisting (at a bare minumum) of the mechanisms of natural selection, sexual selection, genetic drift, and neutral molecular evolution in deep geological time, grounded (at least in part) in theoretical mathematical models of population genetics, depending on multiple sources of heritable phenotypic variation, and supported by inference from multiple sources of empirical evidence, including field and laboratory research in the fields of biochemistry, cell biology, comparative physiology, developmental biology, ecology, ethology, genetics, neurobiology, and physiological ecology.

Note that these two definitions of the principle domains of evolutionary biology correspond roughly to what are sometimes referred to as “macroevolutionary theory” and “microevolutionary theory” (in that order) and do not explicitly mention:

• theories of the origin of life from non-living materials, which are properly the purview of astrophysics, chemistry, and geology, not biology;

• the concept of “adaptation”, which has had a checkered past in evolutionary biology and is facing increasing challenges within the field; and

• teleology, which is almost never mentioned, except for those evolutionary biologists who have thought about it (which, in my experience, are relatively few), who generally assume that resort to teleological explanations in evolutionary biology is unnecessary. Not wrong, just unnecessary (not to mention unproductive as an empirical research hypothesis).

As philosophical concepts, both adaptation and teleology have a very long history, stretching back at least to Plato and Aristotle. However, recent developments in evolutionary theory, including (but not limited to) theories of epigenetics, exaptation, genetic drift/draft, neutral and nearly neutral molecular “drift” in deep evolutionary time, and punctuated equilibrium, have rendered the concept of “adaptation” as an increasingly marginal diversion rather than a central topic in evolutionary biology.

And teleology, rather than being considered “wrong” (when it is considered at all, which is seldom) is now increasingly being incorporated into new theories of “evolved agency”, especially in evolutionary psychology (my own field). I am currently working on a treatise on this latter subject, which I hope to finish before departing this veil of tears and laughter for that undiscovered country from whose bourn no traveller returns.

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Monday, November 16, 2009

The ID Cookie Crumbles...


During this year of celebration of Darwin and evolutionary biology, Intelligent Design (ID) supporters are fond of asserting that a branch of the biological sciences that currently accounts for over 100 regularly published journals (containing over 1000 peer-reviewed scientific reports) per year, over 1000 books published by reputable scientific publishers per year, and involving grant and foundation support amounting to several billion dollars per year is "crumbling", while ID, which accounts for not one peer-reviewed scientific journal and one peer-reviewed book (published over a decade ago) is replacing it.

I can go to Mann Library here at Cornell (the second largest library of biology in the world, comprising over a million books and bound periodicals) and find the equivalent of an entire floor devoted to evolutionary biology. I couldn't carry this month's issues of the various journals devoted to evolutionary biology to the loan desk, even if I used a large laundry basket and made several trips. I have a paltry selection of the most current books on the subject of evolution in my personal library: only 1000+ volumes published in the past ten years or so. If I had unlimited funds, I could buy ten times as many, and still could not keep up with the field.

Virtually every large university in the world has a department of ecology and evolutionary biology. Here at Cornell we have such a department, with almost two dozen professors and dozens of graduate students, and there are at least five other departments at Cornell who number evolutionary biologists among their members. There are almost half a dozen undergraduate and graduate organizations devoted to the scientific aspects of evolutionary biology at Cornell; branches of such societies are found worldwide.

By contrast, there are two tenured professors in the entire world who explicitly support ID, only one of whom is in a department devoted to an empirical science (the other teaches at a theological seminary). Neither of them is currently engaged in empirical research intended to validate ID.

Of the 35+ undergraduate IDEA clubs (a very liberal estimate) that were founded during the heyday of ID (the late 1990s and early 2000s), not one is currently maintaining a website or apparently meeting regularly. And according to Google Trends, interest by the news media in ID has fallen almost to zero since the Kitzmiller v. Dover trial in late 2005, while interest in evolutionary biology is at an all-time high and still increasing with no end in sight.

So, based on the empirical evidence, which is "crumbling", evolutionary biology or ID?

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As always, comments, criticisms, and suggestions are warmly welcomed!

--Allen

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